Preventing Horse-Vehicle Interactions on Assateague

©equus ferus-wild horse photography® & ©karen mclain studio

We need your help

We need the public’s help at Assateague. We need data on any horse-vehicle interactions (fatal, injured, or near-miss) at Assateague Island. We need the exact location, as far back as possible, including near misses. Horses often follow specific trails and paths throughout Assateague, and our ability to pinpoint areas of concern can help prevent future interactions. At the bottom of this page are two maps, one general, the other Google Earth- please feel free to download, add your observation and email us or DM on Facebook.

There has been a great deal of anger and sadness following the recent vehicle strikes on Assateague, and those reactions are completely understandable. When several animals are struck within a short period of time, people naturally look for an explanation, something that changed, something that failed, or something that might have prevented it. I am writing this because I want to help prevent the next horse collision on Assateague Island. I have offered my expertise as a biostatistician to help analyse the collision data and identify patterns that may point toward practical, evidence-based prevention. I care deeply about these horses, but concern alone will not solve the problem. Good data, careful analysis, and realistic mitigation might. I have been in touch with the Assateague Island Alliance.

Susi Solé (N2BHS-M) lost her life on August 13, 2026

Most importantly, science does not support assigning wildlife-vehicle collisions to a single cause. These collisions are multifactorial, involving wildlife behaviour, driver behaviour, visibility, traffic volume, road design, speed, time of day, and surrounding habitat (Gunson et al., 2011; Shilling & Waetjen, 2015).

There is a very real and serious wildlife-vehicle collision problem on Assateague Island. However, this problem is not new. In September 2019, the National Park Service reported that 34 horses had been killed by vehicles and another 17 horses had been injured by vehicles in collisions on the Island since 1982. The Park Service reported that there were 51 documented wildlife-vehicle collisions on the Island since 1982 (National Park Service [NPS], 2019). More recently, in 2021, Moonshadow, a pregnant mare, was killed in a collision on Bayberry Drive. Her foal was seriously injured in the crash. The total number of horses to have been struck by vehicles on the Island was reported to be 53 horses, with 35 of them killed in the resulting collisions (Walburn Viviano, 2021).

The recent cluster of wildlife-vehicle collisions on Assateague Island occurred within a relatively short time frame. In the last six weeks, three horses have been involved in collisions with vehicles on the Island. Two of them were killed in the incidents. Recently, Susi Solé was killed by a vehicle on Bayberry Drive on August 13, 2026. The incident occurred just 13 days after another horse had been killed in a collision with a vehicle on the Island (Sweigart, 2026). In response to recent collisions, the National Park Service (NPS) is investigating additional measures to prevent further incidents involving wildlife and vehicles on the Island. A cluster of wildlife-vehicle collisions does not have to be caused by the same factor(s) that caused the other collisions in the cluster. Therefore, it is not sufficient to conclude that a cluster of wildlife-vehicle collisions has occurred, and then to stop investigating.

©equus ferus-wild horse photography® & ©karen mclain studio

Banning Cars?

As was mentioned previously, there are valid reasons for banning cars from Assateague Island after the recent deaths of several horses. The National Park Service has the authority to ban all vehicles from park areas. However, a permanent, park-wide ban on cars would be far more complicated than simply closing a road or two and then returning to ‘business as usual’ after a temporary period. Already, the National Park Service restricts a variety of activities and uses throughout Assateague Island National Seashore. For example, the over-sand vehicle zone is occasionally closed temporarily for wildlife management purposes. The 36 C.F.R. § 1.5 authority is used to restrict a variety of activities throughout the park.

Furthermore, as previously noted, motor-vehicle travel in national parks is permitted only on park roads and designated routes. Thus, such travel does not constitute an entitlement to travel wherever and whenever one pleases within a national park. Any such ban would likely have a major impact on visitor use of the park and would have to be justified in writing, be subject to public notice, and would likely require additional planning and environmental analysis beyond the authority provided by 36 C.F.R. § 1.5. In fact, the Superintendent’s Compendium for Assateague Island National Seashore states that for major or controversial restrictions on activities, in addition to the authority provided by 36 C.F.R. § 1.5, additional procedures such as formal justification, public notice, planning, and environmental analysis are required.

Therefore, even though a complete car ban would be legally possible, it would be operationally very difficult to carry out. Instead, restrictions on where, when, and how cars could travel in parts of the park where horses are frequently present would be both legally feasible and already consistent with the powers the National Park Service currently exercises at Assateague Island National Seashore. The collision history of horses and cars at Assateague Island would, by itself, not provide a basis for concluding that a complete car ban would be the best solution to the problems that have arisen at the park. Instead, such a history simply raises the question of whether consideration should be given to a variety of other measures that could have the same effect as a complete car ban but which would not have the same impact on visitor use of the park.

©equus ferus-wild horse photography® & ©karen mclain studio

What can be done?

Some examples of such measures include restrictions on nighttime driving by cars, temporary closures of park roads for periods of time when large numbers of horses are using sections of the park’s road system, measures to reduce the speed of approaching cars, measures to control the flow of traffic on park roads and to redistribute the effects of any collisions that might occur, restrictions on the number of cars that could travel on park roads during peak hours of the day, and a ban on cars from certain sections of park roads during periods of time when large numbers of horses are present in those areas of the park. All of these measures are consistent with the mission of the National Park Service and the current management of Assateague Island National Seashore, as outlined in the park’s General Management Plan. The General Management Plan for Assateague Island National Seashore notes that recreation must be resource-compatible. Therefore, while a complete car ban would be legally possible, it would not be the most sensible measure to consider, and instead a number of other, more targeted measures should be considered in order to determine whether or not they would be as effective in reducing the number of collisions between cars and horses as a complete car ban.

Most research on wildlife-vehicle collisions recognises that wildlife-vehicle collisions occur in spatial clusters or hotspots (Gunson et al., 2011). Therefore, the factors explaining wildlife-vehicle collisions in hotspots can vary. For example, factors that explain collisions in one hotspot may not explain collisions in another hotspot of similar wildlife species and road configuration. As such, wildlife-vehicle collision mitigation is best conducted on a site-by-site basis using information specific to each site (Gunson et al., 2011). In addition, research has found that even wildlife-vehicle collision hotspots are not fixed in space over time (Shilling & Waetjen, 2015).

In wildlife-vehicle collision studies, time of year and day of year have also been found to influence collision frequency. Studies have found species-specific patterns in the temporal distribution of wildlife-vehicle collisions. Therefore, for the Assateague horses, a study of wildlife-vehicle collision occurrence would be valuable for identifying the days and times of the year when the greatest number of collisions is expected.

Pollard and Grewar (2020) investigated 4,341 horse-related road incidents in the UK involving ridden or loose horses and found significant spatial and temporal clustering. In terms of factors influencing collision severity, driver behaviour, especially excessive speed, was found to be particularly fatal to horses. Importantly, loose horses (free-roaming ponies) were found to have significantly greater odds of fatal injury than their ridden counterparts.

©equus ferus-wild horse photography® & ©karen mclain studio

Assateague is unique

The information presented above addresses wildlife-vehicle collisions in general and horse-vehicle collisions in general, but not specifically the wild Assateague horses. Some of the information and corresponding data may be applicable to the Assateague horses, but others would not. Several variables have been measured in previous studies of wildlife-vehicle collisions, including speed, visibility, location, time, animal behaviour, and driver behaviour.

1) Well-travelled paths. Wild horses do not move randomly across the landscape. Free-ranging horses establish home ranges and habitually use areas and travel routes in response to water, forage, terrain, season, and other environmental conditions (Hennig et al., 2018; Ostermann-Kelm et al., 2009; Schoenecker et al., 2023). On Assateague specifically, individual bands occupy recognisable home ranges, and long-term research has demonstrated patterned habitat use related to temperature, wind, habitat type, and biting-fly pressure (National Park Service, 2023; Powell et al., 2006). This raises a testable possibility that some road crossings may occur repeatedly where established horse travel routes intersect Bayberry Drive or Route 611.

At Sand Wash Basin, horses approaching a waterhole (yellow circle- Lake Draw) follow familiar, well-worn paths. Although not evident in this photo, steep hills are on either side of the green area near the top. The small pink arrows point to examples of horse trails (though there are many more trails), and the grey is a road/two track

Video by KarenMcLain- we have several GoPros and place them strategically to capture horses coming into water. This video is from Lake Draw, where you can see the horses picking their way down the hill along well-worn horse trails.
©Karen McLain Studio, 2014

We utilised horse trails at Assateague as well
©Karen McLain Studio, 2017.

2) Reducing the speed limit on park roads is not going to result in drivers travelling at lower speeds. Riginos and colleagues (2022) studied the effect of reducing the nighttime speed limit on a section of highway where mule deer frequently cross. They found that, on average, drivers’ speeds were reduced by only about 2%. They concluded that there was little evidence that reducing the posted speed limit on park roads would reduce wildlife-vehicle collisions.

3) Warning systems also have mixed results. Research conducted by Sullivan et al. (2004) in highway migration corridors for mule deer found that temporary, highly visible warning signs with or without flashing amber lights decreased deer-vehicle collisions by about 50% in the treated areas. However, the decrease in collisions was unsustainable because drivers eventually ignored the warning signs. Thus, the warning signs would have to be made highly visible on a temporary basis or be dynamic to be effective.

4) Road-mortality mitigation: Rytwinski et al. (2016) conducted a very large meta-analysis of all sorts of interventions for reducing wildlife roadkill. On average, there was a 40% reduction in wildlife roadkill across interventions, though substantial variation existed among intervention types. For large mammals, the best interventions were fencing and crossing structures. Animal-detection systems seemed to hold a lot of promise but were not yet as effective as, for example, reflectors (which are less expensive to put up).

©equus ferus-wild horse photography® & ©karen mclain studio

National Parks and the Assateague Island Alliance

For those unfamiliar with Assateague, note that the National Park Service (NPS) manages the horses on the Maryland side of the Island as a wildlife population and manages the roads, visitors, etc. within the park. Assateague Island Alliance (AIA) is the official friends group of the park and supports the work of NPS by fundraising, education, volunteers, equipment and conservation programs and activities within the park (NPS, 2015)

The horses are being managed as a wildlife population, and NPS continues to work to reduce food conditioning and habituation of the animals in the park’s developed areas. AIA has been working to help the horses of Assateague by raising $100,000 to replace 222 of the park’s picnic tables with wildlife-resistant food-storage tables. In addition to the work to develop wildlife-resistant food-storage tables, AIA’s funding of the horse-management interns, equipment and visitor-education programs has been a critical component of the Island’s horses’ management as a wildlife population.

©equus ferus-wild horse photography® & ©karen mclain studio

Reasonable mitigation to evaluate could include:

  • Map every collision and near miss event at the exact GPS location to establish hotspots.
  • Document the time of day, lighting, weather, traffic volume, direction of travel on Bayberry Drive and Route 611 and road conditions for each collision or near collision.
  • Measuring actual vehicle speeds, rather than relying only on posted limits.
  • Targeted enforcement and/or traffic-calming measures at locations where speeding or excessive approach speed have been documented.
  • Dynamic or flashing warning systems during identified high-risk periods or in collision hotspots.
  • Nighttime visibility measures (such as improved lighting or increased sight distance) in areas where collisions and near collisions are occurring.
  • Continue to reduce food-conditioning and habituation around developed areas.
  • Recording data on the horses that are struck by vehicles, including the identity of the horses, their bands, their behaviour, and the frequency of their use of the road.
  • Using before-and-after data to establish whether a given intervention does reduce collisions, as opposed to simply being assumed to do so (Rytwinski et al., 2016).
©equus ferus-wild horse photography® & ©karen mclain studio

Using statistics to predict future horse-vehicle interactions

Looking at the most recent incidents first and going into as much detail as possible to improve the data currently being collected. We do not have to treat every horse strike as random. We can build a prediction model that includes season, time of day, traffic volume, and location:


For example, we can look at Bayberry Drive to see if there are any repeats of strikes in the same locations. In addition, we can look at strikes that occur during nighttime hours and at the speeds vehicles are travelling to see whether they are travelling at excessive speeds. Finally, we can look to see which horses are repeatedly occupying the roadways.

In plain English: if the data show that strikes are more common in summer, at night, on Bayberry Drive, and during heavy traffic, the model combines those factors to estimate when collision risk is highest. That is much more useful than simply saying, “another strike will probably happen.” It tells us when and where prevention should be concentrated.

That is how the problem becomes manageable. Three strikes within roughly six weeks are an important signal. It is not yet an explanation. Identify patterns in collisions, measure variables of interest, introduce a few ‘good’ interventions and test for ability to reduce future collisions. Supporting AIA, supporting the National Park Service, and asking for stronger evidence-based collision prevention are not mutually exclusive. The goal should not be to identify somewhere to direct anger.

Our goal here on this blog is to prevent the next horse collision on Assateague Island. Thank you for your time and consideration. Together we can.

Part of the compiled database

Dr Meredith Hudes-Lowder, Biostatistician
August 16, 2026


References

Assateague Island Alliance. (n.d.). Donations and Protect initiatives. Assateague Island Alliance.

Gunson, K. E., Mountrakis, G., & Quackenbush, L. J. (2011). Spatial wildlife-vehicle collision models: A review of current work and its application to transportation mitigation projects. Journal of Environmental Management, 92(4), 1074–1082. https://doi.org/10.1016/j.jenvman.2010.11.027

Hennig, J. D., Beck, J. L., & Scasta, J. D. (2018). Spatial ecology observations from feral horses equipped with global positioning system transmitters. Human–Wildlife Interactions, 12(1), 75–84. https://doi.org/10.26077/z9cn-4h37

King, S. R. B., & Schoenecker, K. A. (2022). Application of tail transmitters for tracking feral horses as an alternative to radio collars. Wildlife Society Bulletin, 46(4), e1338. https://doi.org/10.1002/wsb.1338

National Park Service. (2015). Join our friends: Assateague Island Alliance. Assateague Island National Seashore.

National Park Service. (2019, September 30). Assateague Island National Seashore wild horse fatality. Assateague Island National Seashore.

National Park Service. (2022, May 2). Relocation of an aggressive and highly food-conditioned horse from Assateague Island National Seashore to the Cleveland Amory Black Beauty Ranch. Assateague Island National Seashore.

National Park Service. (2023, May 16). Assateague’s wild horses. Assateague Island National Seashore. https://www.nps.gov/asis/learn/nature/horses.htm

National Park Service. (2026). 2026 annual horse population update for Assateague Island National Seashore. Assateague Island National Seashore.

Ostermann-Kelm, S. D., Atwill, E. A., Rubin, E. S., Hendrickson, L. E., & Boyce, W. M. (2009). Impacts of feral horses on a desert environment. BMC Ecology, 9, 22. https://doi.org/10.1186/1472-6785-9-22

Pollard, D., & Grewar, J. D. (2020). Equestrian road safety in the United Kingdom: Factors associated with collisions and horse fatalities. Animals, 10(12), 2403. https://doi.org/10.3390/ani10122403

Powell, D. M., Danze, D. E., & Gwinn, M. A. (2006). Predictors of biting fly harassment and its impact on habitat use by feral horses (Equus caballus) on a barrier island. Journal of Ethology, 24(2), 147–154. https://doi.org/10.1007/s10164-005-0174-2

S. Raymond, A. L. W. Schwartz, R. J. Thomas, E. Chadwick, S. E. Perkins, “Temporal patterns of wildlife roadkill in the UK,” PLOS ONE, vol. 16, no. 10, p. e0258083, 2021. https://doi.org/10.1371/journal.pone.0258083

C. Riginos, E. Fairbank, E. Hansen, J. Kolek, and M. P. Huijser. 2022. Reduced speed limit is ineffective for mitigating the effects of roads on ungulates. Conservation Science and Practice 4(3):e618. https://doi.org/10.1111/csp2.618

T. Rytwinski, K. Soanes, J. A. G. Jaeger, L. Fahrig, C. S. Findlay, J. Houlahan, R. van der Ree, and E. A. van der Grift. 2016. How effective is road mitigation at reducing road-kill? A meta-analysis. PLOS ONE 11(11): e0166941. https://doi.org/10.1371/journal.pone.0166941

Schoenecker, K. A., Esmaeili, S., & King, S. R. B. (2023). Seasonal resource selection and movement ecology of free-ranging horses in the western United States. The Journal of Wildlife Management, 87(2), e22341. https://doi.org/10.1002/jwmg.22341

Shilling, F. M., & Waetjen, D. P. (2015). Wildlife-vehicle collision hotspots at US highway extents: Scale and data source effects. Nature Conservation, 11, 41–60. https://doi.org/10.3897/natureconservation.11.4438

Sullivan, T. L., Williams, A. F., Messmer, T. A., Hellinga, L. A., & Kyrychenko, S. Y. (2004). Effectiveness of temporary warning signs in reducing deer-vehicle collisions during mule deer migrations. Wildlife Society Bulletin, 32(3), 907–915.

Oscar Sweigart, “Three horses struck, two fatally within weeks on Assateague Island,” WBOC, 14 August 2026.

Walburn Viviano, M. (2021, July 11). Assateague pony killed, foal injured by hit & run driver. Chesapeake Bay Magazine/Bay Bulletin.


Maps to download
Indicate the location of any horse/car interactions with an X and email merry@equusferus.com or DM through our Facebook page. Please include (if known) the horse’s name, date, and time of day. Right-click the image, select “Save image as,” and save it to your hard drive. If you have trouble, we can email you a copy.

Gaia Map
Google Earth Map
©equus ferus-wild horse photography® & ©karen mclain studio
©equus ferus-wild horse photography® & ©karen mclain studio

The End

The Tobiano Paints

 THE TOBIANO PAINTS

Disclaimer: The mustang photographs on this blog post are presented without genetic testing; we do not know the actual chromosomal make-up of the mustangs. We rely solely upon the horse’s phenotype, or how they appear physically: coat colour, white markings, eye color, mane, and tail colour.

Mammalian Pigmentation

The colors found in mammalian hair, skin, irises, and some internal organs is produced by the pigment melanin. Melanin appears as colored granules in these pigmented cells and occurs in two forms, eumelanin, and phaeomelanin. Eumelanin is responsible for brown and black colour, and phaeomelanin is responsible for reds and yellows (Bailey & Brooks, 2013).
Black tobiano (muddy) (Black Hills Wild Horse SanctuarySouth Dakota) © Equus ferus- Wild Horse Photography ™ © Karen McLain

The absence of melanin will appear as white in mammals. The lack of color typically associated with Paint Horses is caused by the inability of these cells to produce the base colors from birth. The color loss results in large patches of white against a base coat of any colour (bay, chestnut, roan, grey, dun, champagne, silver dapple, palomino, brown, black, etc). Additionally, the white colour has pink skin beneath. 

Assorted tobiano horses (Black Hills Wild Horse SanctuarySouth Dakota) © Equus ferus- Wild Horse Photography ™ © Karen McLain

There are other forms of white colouration in equines; for example gray or roan. Grey horse color is caused by the failure of melanocytes over time, so the hair starts normally pigmented but loses the ability to maintain the pigment so the horse eventually turns white(Sponenberg, 2009). Roan horses are roan from birth although they are often not recognizable until after the foal coat has shed. Roan horses typically retain the base color on their head, legs, mane, and tail. The skin beneath grey and roan horses is dark and these colors are not actual colours, but rather modifiers that act upon a base coat (Gower, 2016). 

Black tobiano demonstrating the “shield” (Black Hills Wild Horse Sanctuary, South Dakota) © Equus ferus Wild Horse Photography © Karen McLain 
A tobiano horse may also be roan, appaloosa, sabinos, overo, or any other pattern as the tobiano pattern is not a mutually exclusive coat patterns.
Bay Roan tobiano   (Great Desert Basin, Utah) © Equus ferus- Wild Horse Photography ™ © Karen McLain
Roan tobiano foal, palomino tobiano mare  (Great Desert Basin, Utah) © Equus ferus- Wild Horse Photography ™ © Karen McLain

Equine chromosomes
The genes for four white coat patterns: roan (RN), sabinos (SB1), dominant white (W), and tobianos (TO) are located on the KIT gene.The KIT gene is responsible for sending instructions through cells that allow the cell to make specific proteins. The KIT proteins are found on the cell membrane where another protein called a “stem cell factor” binds to the KIT protein. When bound together, they activate the KIT protein, which in turn, activates other proteins within the cell. These proteins serve a variety of functions in mammalian cells such as growth, development, migration, and production of certain cell types such as interstitial gastrointestinal cells and melanocytes (Haase, Jude, Brooks, & Leeb, 2008).
Equine Chromosome #3
The KIT gene is located on the fourth chromosome at the 12 position and located very close is the ECA3 gene. The ECA3, or the third equine chromosome is the location of the chromosomal mutation responsible for tobianos. Although the KIT gene remains normal in these horses, the third chromosome has an area in the gene that has flipped. Approximately one-third the length of the chromosome is an area that is an exact mirror image in tobianos horses. Because the chromosomal inversion is adjacent to the KIT gene, it affects the KIT protein synthesis, and the cells cannot produce melanocytes- so the horse has areas of white. The test for tobianos examines the chromosome and looks for a ‘break’ (telomeric or centromeric) at the positions 13 and 21 on the third chromosome– this serves as an indication they separated and inverted during replication- the horse is genetically a tobianos (Bailey & Brooks, 2009).
The gene for tobiano horses is autosomal dominant. This means to be tobiano, a foal must have at least one tobiano parent, but they also may have two tobiano parents. If one parent is a tobiano, it does not matter what colour or pattern the other parent appears; the foal will be tobiano. If a horse matches the criteria for a tobiano, it is likely the horse has the genetic background of a tobiano, although there may be some mixing of other patterns (Gower, 2016). The patterns are not inherited exactly, however, the proportion of white to colour is inherited. In other words a horse with a lot of white will have offspring with a lot of white but this depends upon the other parent. Interestingly, the study by Woolf (1990) discovered male horses and those with chestnut coats have more white than female or bay horses. The researcher also noted that the inheritance of white leg markings and facial markings is multifactorial; there are many genes involved in the appearance of white markings (Woolf, 1990).

History:

Bay tobiano mare  (McCullough Peaks, Wyoming) © Equus ferus- Wild Horse Photography ™ © Karen McLain

The Tobiano paint pattern was named for General Tobías from Brazil. The General brought the paint horses to Argentina in the mid-1800’s. Before their arrival, tobiano horses were rare and had been grouped with other spotted-type horses. After General Tobías’ arrival, they were renamed after the general and placed into a unique paint coat classification(Kerson, 2015; Sponenberg, 2009)
Tobiano Characteristics:
Bay tobiano mare (McCullough Peaks, Wyoming) © Equus ferus- Wild Horse Photography ™ © Karen McLain
The tobiano is defined by several coat characteristics. As with all horses, unless genetically tested, we evaluate the coat pattern by the phenotype or the horses’ physical appearance. As a general rule, tobiano horses have the following characteristics (there are always exceptions to these rules):

  1. White cross the spine somewhere between the ears and the tail (Gower, 2016; Sponenberg, 2009)
  2. The body white appears to travel down in a vertical fashion (Gower, 2016)
  3. The edges of the white areas tend to be crisp and well-defined (Sponenberg, 2009)
  4. Legs are white and the edge of the socks/stockings is irregular (Gower, 2016; Sponenberg, 2009)
  5. Most tobiano have dark eyes although some tobiano horses have blue eyes  (Sponenberg, 2009)
  6.  Most tobianos have white areas within the mane and tail, this gives the appearance of a bicoloured tail, a trait usually seen only in tobiano horses. (Sponenberg, 2009)
  7. The predominantly solid coloured heads of tobianos are generally conservatively marked: thin blazes, simple stars (Sponenberg, 2009)

 Some tobianos have very little white

Black tobiano (McCullough Peaks, Wyoming) © Equus ferus- Wild Horse Photography ™ © Karen McLain
Minimally marked black tobiano stallion -note the bicoloured tail (McCullough Peaks, Wyoming) © Equus ferus- Wild Horse Photography ™ © Karen McLain
Bay tobiano stallion (McCullough Peaks, Wyoming) © Equus ferus- Wild Horse Photography ™ © Karen McLain
Black tobiano stallion (McCullough Peaks, Wyoming) © Equus ferus- Wild Horse Photography ™ © Karen McLain
Bay tobiano stallion (McCullough Peaks, Wyoming) © Equus ferus- Wild Horse Photography ™ © Karen McLain
Some tobiano horses have very little base colour but they tend retain normally coloured heads even when extensively white.
Minimally marked light bay chestnut tobiano (Black Hills Wild Horse Sanctuary, South Dakota) © Equus ferus- Wild Horse Photography ™ © Karen McLain
Light bay tobiano (note the minimal blaze) (Black Hills Wild Horse SanctuarySouth Dakota) © Equus ferus- Wild Horse Photography ™ © Karen McLain
Extensively white black marked tobiano (Black Hills Wild Horse SanctuarySouth Dakota) © Equus ferus- Wild Horse Photography ™ © Karen McLain
Some tobiano horses have marks within the white areas and there may be some bleeding of colour between the base colour and the white areas. The smaller spots in the white areas are called ink spots, bear tracks, cat’s paws. The areas of darker colour encroaching on the white areas are referred to as halos. There may also be some roaning at the edge of colour and white. This is a historical link between these markings and homozygosity. No genetic link has been found, however anecdotally, homozygous horses often present with these marking whilst heterozygous generally do not show these markings. 

Cat’s paws & halo effect (San Wash Basin, Colorado) © Equus ferus- Wild Horse Photography ™ © Karen McLain

References
Bailey, E., & Brooks, S. (2009). Method for screening for a tobiano coat color genotype  #USPatent 8101354 B2.
Bailey, E., & Brooks, S. (2013). Horse Genetics (2nd ed.). Boston, Massachusetts: CABI.
Gower, J. (2016). Horse Color Explained: A Breeder’s Perspective. Brattleboro, Vermont: Echo Point Books & Media, Inc.
Haase, B., Jude, R., Brooks, S. A., & Leeb, T. (2008). An equine chromosome 3 inversion is associated with the tobiano spotting pattern in German horse breeds. 
Animal Genetics, 39(3), 306-309. doi:10.1111/j.1365-2052.2008.01715.x
Kerson, N. (2015). What Color is that?  A quick guide to horse color identification: Nancy Kerson- Self Published
Sponenberg, D. (2009). Equine Color Genetics (3rd ed.). Ames, Iowa: Wiley-Blackwell.
Woolf, C. M. (1990). Multifactorial inheritance of common white markings in the Arabian horse. J Hered, 81(4), 250-256.

A special thanks to Nancy Kerson for her brilliant book “What Color is that? A quick guide to horse color identification” and to the Black Hills Wild Horse Sanctuary. A worth sanctuary for wild horses.

Assateague Island National Park 2015

Hello Everyone!!!
We’re back at Assateague Island NP for a long weekend of photography.  Karen McLain has already been here for over a week as an “Artist-in-Residence”. This is a prestigious appointment in which an artist, often a painter, is invited to spend a fortnight at a location and uses the opportunity to hone their artistic skill in a new environment and they often teach classes as well. 


I joined Karen today after driving down the coast from the Hudson Valley in New York. We shot a few of the ponies, although the conditions were less than optimal with an intermittent light mist falling. We broke early and had a wonderful dinner at a restaurant in Salisbury, Maryland called “Brew River”- the specialty was crab cakes and they did the cakes justice.  The oysters on the half-shell were amazing and we thoroughly enjoyed this dining experience.



It was also nice to see a horse that had been a foal, when we were last here in 2012. One of the things I particularly enjoy is seeing foals grow up and get bands of their own or give birth to their own foals. It lends a more personal aspect to the photography.
Foal 2015- Today
Foal 2012
Same foal today 2015


Tomorrow we are up early and plan to have a full day in the park. Karen is teaching a class for the Assateague Island Alliance on Saturday so we will prepare for that- a local paper did a nice segment of Karen’s residency here . 


As always, continue sharing Equus ferus’ photos! We will be posting live from the park.